Greenhouses: Welwitschia-annex
The two subspecies of Welwitschia
mirabilis in Berlin-Dahlem
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Old stock of Welwitschia mirabilis
with both subspecies in the nursery in summer 2001. |
Besides the old and young Welwitschia plants in the display
glasshouse, thirteen plants of old stock are growing in the Welwitschia
house in the nursery (not open to the public for reasons of limited
space). All were grown from seed. Trunk diameters measured in
September 2000 were between 15 and 45 cm and leaf widths between 16
and 61 cm. Of the 13 plants, eight are producing male cones, four have
at least once produced female cones, three have not flowered up to now
(year 2001). A constant soil temperature of c. 26°C maintained in
the glasshouse in the nursery has proved to be highly beneficial for
successful growth and profuse flowering of our Welwitschia
mirabilis. The Berlin-Dahlem horticulturist working with the Welwitschia
collection at Berlin-Dahlem since 1964, Mr. Kurt Swiatkowiak was first
to observe differences in flowering period and in male cone characters
between individual plants among the simultaneously flowering male
welwitschias of the old stock. His repeated observations on peduncle
length, shape, and colour of the male cones led the author to attempt
a closer look some years ago. However, the lack of exact documentation
of origin of the plants of the old and presumably heterogeneous stock
was a major impediment for a systematic approach to such a study. The
observations suggested that the old Welwitschia stock at Berlin-Dahlem
was composed of two groups of plants, provisionally denominated as "group
1" and "group 2". Only in 1999 and 2000, a search for
more data on the origin of the seeds received from the Botanical
gardens of Coimbra (seed origin Angola) and Kirstenbosch (seed origin
Namibia) and a comparison with plants illustrated in literature and
with herbarium material of known origin finally permitted to present
results that throw new light on this often studied plant. A
new
subspecies, Welwitschia mirabilis subsp. namibiana
was published in December 2001 in the journal Willdenowia.
The Berlin-Dahlem plants of "group 1" with reddish brown
male cones agree with material from Angola and belong to Welwitschia
mirabilis subsp. mirabilis, the tardily flowering plants
with glaucous-green male cones of "group 2" are of Namibian
origin and belong to the new subspecies Welwitschia mirabilis
subsp. namibiana. The relevant differences, however, are not
so much found in the colour of the cones but in details of the bract
shape of the male cones.
Representative plants with male cones and details of male cones of
both subspecies are illustrated side by side below: |
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Flowering male Welwitschia
mirabilis subsp. mirabilis (plant nr. 2) in August 2000. |
Flowering male Welwitschia
mirabilis subsp. namibiana (plant nr. 6) in August 2000. |
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Male cones of Welwitschia
mirabilis subsp. mirabilis (plant nr. 2) in August 2000. |
Male cones of Welwitschia
mirabilis subsp. namibiana (plant nr. 6) in August 2000. |
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Male cones of Welwitschia
mirabilis subsp. mirabilis (plant nr. 12) in 1981. |
Male cones of Welwitschia
mirabilis subsp. namibiana (plant nr. 3) in September 2001. |
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Detail of male cone and flower of
Welwitschia mirabilis subsp. mirabilis (plant nr. 9). |
Detail of male cone and flower of
Welwitschia mirabilis subsp. namibiana (plant nr. 6). |
[Text: Beat Ernst Leuenberger] |
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