Willdenowia – Annals of the Botanic Garden and
Botanical Museum Berlin-Dahlem
ISSN 0511-9618
© 2004 BGBM Berlin-Dahlem.
Businsky, R. 2004: A revision of the Asian Pinus
subsection Strobus (Pinaceae).
Willdenowia 34: 209-257.
doi:10.3372/wi.34.34120
Abstract
The Asian taxa of Pinus subsect. Strobus are
revised, based on available type and other authentic material as
well as on extensive field studies, including populations at the
type localities. P. fenzeliana is neotypified and transferred to
P. subsect. Flexiles. Statistical evaluation of
ample material revealed that the original description of P.
kwangtungensis is based on heterogeneous material, including a
population from Hainan Island, which is here distinguished and
described as the new species P. orthophylla. The continental
P. kwangtungensis s. str., which is morphologically very
close to P. wangii, is recognized as a subspecies of the
latter, besides subsp. varifolia. An isolated population in
N Vietnam, near the Laotian border, hitherto also considered to
belong to P. kwangtungensis, is described as the new species
P. eremitana. The Japanese P. parviflora, with two
subspecies, is found to be closely related to P. wangii and P.
eremitana. The imperfectly known Taiwanese P. uyematsui,
mostly included in the other Taiwanese endemic P. morrisonicola,
is maintained as a separate species similar to the Vietnamese P.
dalatensis, and a hybrid between the two Taiwanese species is
described as the nothospecies P. ×hayatana. The
E Himalayan P. bhutanica is recognized as a subspecies of
P. wallichiana. A key to all members of P. subsect.
Strobus, comprising nine Eurasian and six American species,
and a revised classification of the Eurasian species in two series
and three subseries, one series and two subseries here described as
new, are provided.
Full-text article | Appendix 1+2: Herbarium material examined | Statistical and biometrical data